Networks receiving negative-derivative feedback in multiple Fourier components and maintenance of multiple bumps of activity. A, B, Example networks with negative-derivative feedback in multiple Fourier components (A). The synaptic connectivity profiles for this example were Gaussian shaped (B). C, D, The presence of negative-derivative feedback in higher order Fourier components permits the maintenance of narrowly tuned patterns of activity (C) and multiple bumps of activity (D). Data in C and D are shown at 3 s into the delay period.